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    <meta content="Jordan, Gregory J." name="eprints.creators_name" />
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<meta content="A new genus and species (Caryophylloflora paleogenica genus &amp; species nova G.J.Jord. &amp; Macphail) are proposed for a fossil inflorescence found in Middle-Late Eocene sediments at Locharbour, north-eastern Tasmania, Australia. A parsimony analysis of 75 extant species of the order Caryophyllales and five outgroups placed the fossil within Caryophyllaceae, either subfamily Alsinoideae or Caryophylloideae. The analysis used molecular (rbcL and/or matK), morphological and anatomical data for the extant species, and morphological data for the fossil. Tests on extant species imply that the placement of the fossil should be convincing. The fossil appears to be of a lineage distinct from any extant Australian Caryophyllaceae. In situ pollen are consistent with the form species, Periporopollenites polyoratus. This relatively simple pollen type first appears in Australia and New Zealand in the Late Cretaceous, the oldest known record of the Caryophyllaceae. The last appearance of P. polyoratus in Australia is in the Oligocene, and extant Australian members of the Caryophyllaceae may have evolved from species that dispersed from elsewhere during the Neogene or Quaternary." name="eprints.abstract" />
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<meta content="BAILLIE, P. W. 1989. Jurassic-Cainozoic. In C. F. Burrett and E. L. Martin [eds.], Geology and mineral resources of Tasmania, 339-409. Geological Society of Australia, Melbourne, Australia.
BARNES, R. W., AND G. J. JORDAN. 2000. Eucryphia (Cunoniaceae) reproductive and leaf macrofossils from Australian Cainozoic sediments. Australian Systematic Botany 13: 373-394.
BEHNKE, H-D. 1994. Sieve-element plastids: their significance for the evolution and systematics of the order. In H-D. Behnke and J. T. Mabry [eds.], Caryophyllales: evolution and systematics, 87-122. Springer Verlag, Berlin, Germany.
BITTRICH, V. 1993. Caryophyllaceae. In K. Kubitzki, J. G. Rohwer and V. Bittrich [eds.], The families and genera of vascular plants, vol II. Flowering plants, Dicotyledons, Magnoliid, Hamamelid and Caryophyllid Families, 206-236. Springer-Verlag, Berlin, Germany. 
CARPENTER, R. J., R. S. HILL, AND G. J. JORDAN. 1994. Cenozoic vegetation in Tasmania. In R. S. Hill [ed.], History of the Australian vegetation: Cretaceous to Recent, 276-298. Cambridge University Press, Cambridge, UK.
CHANDLER, E. M. 1961. Flora of the Lower Headon Beds of Hampshire and the Isle of Wight. Bulletin of the British Museum of Natural History: Geology 5: 91-158.
CHANDLER, E. M. 1963. The lower Tertiary floras of South England. III. Flora of the Bournemouth Beds; the Boscombe, and the Highcliff Sands, The British Museum, London, UK.
CLEMENTS, J. S., J. T. MABRY, H. WYLER, AND A. S. DRIEDING. 1994. Chemical review and evolutionary significance of betalaines. In H-D. Behnke and J. T. Mabry, [eds.], Caryophyllales, evolution and systematics, 247-262. Springer Verlag, Berlin, Germany.
CORREA, M. N. 1984. Flora Patagonica. Instituto Nacional de Tecnologia Agropecuaria. Buenos Aires, Argentina.
CRONQUIST, A. 1981. An integrated system of classification of flowering plants. Columbia University Press, New York, New York, USA.
CUENOD, P., V. SAVOLAINEN, L. W. CHATROU, M. POWELL, R. E. J. GRAYER, AND M. W. CHASE. 2002. Molecular phylogenetics of Caryophyllales based on nuclear 18s rDNA and plastid rbcL, atpB, and matK DNA sequences. American Journal of Botany 89: 132–144
DAVIS, P. H. 1967. Flora of Turkey and the East Aegean Islands, vol. 2. Edinburgh University Press, Edinburgh, UK.
DILCHER, D. L. 2000. Toward a new synthesis: Major evolutionary trends in the angiosperm fossil record. Proceedings of the Natonal Academy of Sciences 97: 7030-7036.
DOWNIE, S. R., AND D. PALMER. 1994. A chloroplast DNA phylogeny of the Caryophyllales based on structural and inverted repeat restriction site analysis. Systematic Botany 19: 236-252.
DYER, R. A. 1975. The genera of southern African flowering plants, vol. 1: Dicotyledons. Department of Agriculture Technical Services, Pretoria, South Africa
ELIASSON, U. 1993. 35A, Phytolaccaceae, Flora of Ecuador, 46. University of Goteborg Press, Goteborg, Sweden.
HEUSSER, C. J., 1971. Pollen and Spores of Chile. Modern Types of the Pteridophyta, Gymnospermae, and Angiospermae. University of Arizona Press, Tucson, USA.
HILL, R. S. 1989 New species of Phyllocladus (Podocarpaceae) macrofossils from south eastern Australia. Alcheringa 13: 193-208.
HILL, R. S., AND R. J. CARPENTER. 1991. Evolution of Acmopyle and Dacrycarpus (Podocarpaceae) foliage as inferred from macrofossils in south-eastern Australia. Australian Systematic Botany 4: 449-479.
HILL, R. S., AND D. C. CHRISTOPHEL. 1988. Tertiary leaves of the tribe Banksieae (Proteaceae) from southeastern Australia. Botanical Journal of the Linnean Society 97: 205-27.
HILL, R. S., AND M. S. POLE. 1992. Leaf and shoot morphology of extant Afrocarpus, Nageia and Retrophyllum (Podocarpaceae) species, and species with similar leaf arrangement from Tertiary sediments in Australasia. Australian Systematic Botany 5: 337-58.
HNATIUK, R. J. 1990. Census of Australian vascular plants. Australian Government Publishing Service, Canberra, Australia, 650 pp.
HOFMANN, U. 1994. Flower morphology and ontogeny. In H -D. Behnke and J. T. Mabry [eds.], Caryophyllales, evolution and systematics, 123-166. Springer Verlag, Berlin, Germany.
HEUSSER, C. J. 1971. Pollen and spores of Chile. University of Arizona Press, Tucson, Arizona, USA.
JORDAN, G. J. 2001. An investigation of long-distance dispersal based on species native to both Tasmania and New Zealand. Australian Journal of Botany 49 333-340.
JORDAN, G. J., AND R. S. HILL. 2000. The phylogenetic affinities of Nothofagus leaf fossils based on combined molecular and morphological data. International Journal of Plant Science 160: 1177-1188.
KÄLLERSJÖ, M., J. S. FARRIS, A. G. KLUGE, AND C. BULT. 1992. Skewness and permutation. Cladistics 8: 275-287.
KUBITZKI, K., J. G. ROHWER, AND V. BITTRICH. 1993. The families and genera of vascular plants, vol II. Flowering plants, Dicotyledons, Magnoliid, Hamamelid and Caryophyllid Families. Springer-Verlag, Berlin, Germany.
MACPHAIL, M. K. 1997. Late Neogene climates in Australia: fossil pollen- and spore-based estimates in retrospect and prospect, Australian Journal of Botany 45: 425-464.
MACPHAIL, M. K., N. ALLEY, E. M. TRUSWELL, AND I.R. SLUITER. 1994. Early Tertiary vegetation: evidence from pollen and spores. In R. S. Hill, [ed.], History of the Australian vegetation: Cretaceous to Recent, 189-261. Cambridge University Press, Cambridge, UK.
MANHART, J. R., AND J. H. RETTIG. 1994. Gene sequence data. In H-D. Behnke and J. T. Mabry [eds.], Caryophyllales: evolution and systematics, 235-246. Springer Verlag, Berlin, Germany.
MANLY, W. C., AND C. R. HUTCHINS. 1980. A flora of New Mexico. J. Cramer, Vaduz, Germany.
MOAR, N. T. 1993. Pollen grains of New Zealand dicotyledonous plants. Manaaki Whenua Press, Lincoln, New Zealand.
MULLER, J. 1981. Fossil pollen records of extant angiosperms. Botanical Review 47: 1-142.
NOTT, J. F., AND J. A. K. OWEN. 1992. An Oligocene palynoflora from the middle Shoalhaven catchment N.S.W. and the Tertiary evolution of flora and climate in the southeast Australian highlands. Palaeogeography, Palaeoclimatology, Palaeoecology 95: 135-151.
NOWICKE, J. W. 1994. Pollen morphology and exine ultrastructure. In H-D. Behnke and J. T. Mabry [eds.], Caryophyllales: evolution and systematics, 168-221. Springer Verlag, Berlin, Germany.
NOWICKE, J. W. 1996. Pollen morphology, exine structure and the relationships of Basellaceae and Didieraceae to Portulacaceae. Systematic Botany 21: 187-208,
NOWICKE, J. W., AND J. J. SKVARLA. 1979. Pollen morphology: the potential influence in higher order systematics. Annals of the Missouri Botanic Gardens 66: 633-700.
OWEN, J. A. K. 1988. Miocene palynomorph assemblages from Kiandra, New South Wales. Alcheringa 12: 269-297.
PRENDINI, L. 2001. Species or supraspecific taxa as terminals in cladistic analysis? Groundplans versus exemplars revisited. Systematic Biology 50: 290–300.
PUNT, W., AND P. P. HÖEN. 1995. Caryophyllaceae. Review of Palaeobotany and Palynology 88: 82-272.
RAINE, J. I. 1984. Outline of a palynological zonation of Cretaceous to Paleogene terrestrial sediments in West Coast region, South Island, New Zealand. New Zealand Geological Survey Report 109: 1-82. 
RODMAN, J. E. 1994. Cladistic and phenetic studies. In H-D. Behnke and J. T. Mabry [eds.], Caryophyllales: evolution and systematics, 279-301. Springer Verlag, Berlin, Germany.
ROLES, S. J. 1957. Flora of the British Isles, illustrations, part 1. Cambridge University Press, Cambridge, UK.
ROSS-CRAIG, S. 1951. Drawings of the British Flora. Vol. 5. Caryophyllaceae. G. Bell and Sons, London, Great Britain.
SAVOLAINEN, V., et al.. 2000. Phylogeny of the eudicots: a nearly complete familial analysis based on rbcL gene sequences. Kew Bulletin 555: 257-309.
SIMMONS, M. P., AND H. OCHOTERENA 2000 Gaps as characters in sequence-based phylogenetic analyses. Systematic Biology 49: 369-381
SMISSEN, R. D., J. C. CLEMENT, P. J. GARNOCK-JONES, AND G. K. CHAMBERS 2002. Subfamilial relationships within Caryophyllaceae as inferred from 5' ndhF sequences. American Journal of Botany 89: 1336–1341.
STOVER, L. E., AND A. D. PARTRIDGE. 1973. Tertiary and Late Cretaceous spores and pollen from the Gippsland Basin, south-eastern Australia. Proceedings of the Royal Society of Victoria 85: 237-86.
SWOFFORD, D. L. 2000. PAUP*: phylogenetic analysis using parsimony (*and other methods), version 4. Sinauer Associates, Sunderland, Massachusetts, USA.
TRUSWELL, E. M., I. R. SLUITER, AND W. K. HARRIS. 1985. Palynology of the Oligocene-Miocene sequence in the Oakvale-1 corehole, western Murray Basin, South Australia. BMR Journal of Australian Geology &amp; Geophysics 9: 267-295.
TULIP, J. R., G. TAYLOR, AND E. M. TRUSWELL. 1982. Palynology of Tertiary Lake Bunyan, Cooma, New South Wales. BMR Journal of Australian Geology &amp; Geophysics 7: 255-268.
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<meta content="A new genus and species (Caryophylloflora paleogenica genus &amp; species nova G.J.Jord. &amp; Macphail) are proposed for a fossil inflorescence found in Middle-Late Eocene sediments at Locharbour, north-eastern Tasmania, Australia. A parsimony analysis of 75 extant species of the order Caryophyllales and five outgroups placed the fossil within Caryophyllaceae, either subfamily Alsinoideae or Caryophylloideae. The analysis used molecular (rbcL and/or matK), morphological and anatomical data for the extant species, and morphological data for the fossil. Tests on extant species imply that the placement of the fossil should be convincing. The fossil appears to be of a lineage distinct from any extant Australian Caryophyllaceae. In situ pollen are consistent with the form species, Periporopollenites polyoratus. This relatively simple pollen type first appears in Australia and New Zealand in the Late Cretaceous, the oldest known record of the Caryophyllaceae. The last appearance of P. polyoratus in Australia is in the Oligocene, and extant Australian members of the Caryophyllaceae may have evolved from species that dispersed from elsewhere during the Neogene or Quaternary." name="DC.description" />
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    <h1 class="ep_tm_pagetitle">A Middle-Late Eocene inflorescence of Caryophyllaceae from Tasmania, Australia</h1>
    <p style="margin-bottom: 1em" class="not_ep_block"><span class="person_name">Jordan, Gregory J.</span> and <span class="person_name">Macphail, Michael K.</span> (2003) <xhtml:em>A Middle-Late Eocene inflorescence of Caryophyllaceae from Tasmania, Australia.</xhtml:em> American Journal of Botany, 90 (5). pp. 761-768.</p><p style="margin-bottom: 1em" class="not_ep_block"></p><table style="margin-bottom: 1em" class="not_ep_block"><tr><td valign="top" style="text-align:center"><a href="http://eprints.utas.edu.au/1727/1/Caryophyllaceae.pdf"><img alt="[img]" src="http://eprints.utas.edu.au/style/images/fileicons/application_pdf.png" border="0" class="ep_doc_icon" /></a></td><td valign="top"><a href="http://eprints.utas.edu.au/1727/1/Caryophyllaceae.pdf"><span class="ep_document_citation">PDF</span></a> - Full text restricted - Requires a PDF viewer<br />1417Kb</td><td><form method="get" accept-charset="utf-8" action="http://eprints.utas.edu.au/cgi/request_doc"><input value="2223" name="docid" accept-charset="utf-8" type="hidden" /><div class=""><input value="Request a copy" name="_action_null" class="ep_form_action_button" onclick="return EPJS_button_pushed( '_action_null' )" type="submit" /> </div></form></td></tr></table><p style="margin-bottom: 1em" class="not_ep_block">Official URL: <a href="http://www.amjbot.org/">http://www.amjbot.org/</a></p><div class="not_ep_block"><h2>Abstract</h2><p style="padding-bottom: 16px; text-align: left; margin: 1em auto 0em auto">A new genus and species (Caryophylloflora paleogenica genus &amp; species nova G.J.Jord. &amp; Macphail) are proposed for a fossil inflorescence found in Middle-Late Eocene sediments at Locharbour, north-eastern Tasmania, Australia. A parsimony analysis of 75 extant species of the order Caryophyllales and five outgroups placed the fossil within Caryophyllaceae, either subfamily Alsinoideae or Caryophylloideae. The analysis used molecular (rbcL and/or matK), morphological and anatomical data for the extant species, and morphological data for the fossil. Tests on extant species imply that the placement of the fossil should be convincing. The fossil appears to be of a lineage distinct from any extant Australian Caryophyllaceae. In situ pollen are consistent with the form species, Periporopollenites polyoratus. This relatively simple pollen type first appears in Australia and New Zealand in the Late Cretaceous, the oldest known record of the Caryophyllaceae. The last appearance of P. polyoratus in Australia is in the Oligocene, and extant Australian members of the Caryophyllaceae may have evolved from species that dispersed from elsewhere during the Neogene or Quaternary.</p></div><table style="margin-bottom: 1em" border="0" cellpadding="3" class="not_ep_block"><tr><th valign="top" class="ep_row">Item Type:</th><td valign="top" class="ep_row">Article</td></tr><tr><th valign="top" class="ep_row">Additional Information:</th><td valign="top" class="ep_row">The definitive version is available at http://www.amjbot.org/cgi/content/abstract/90/5/761</td></tr><tr><th valign="top" class="ep_row">Keywords:</th><td valign="top" class="ep_row">Antarctica; Australia; Caryophyllaceae; Centrospermae; Eocene; fossil; Paleogene; phylogeny; pollen</td></tr><tr><th valign="top" class="ep_row">Subjects:</th><td valign="top" class="ep_row"><a href="http://eprints.utas.edu.au/view/subjects/260110.html">260000 Earth Sciences &gt; 260100 Geology &gt; 260110 Biostratigraphy</a><br /><a href="http://eprints.utas.edu.au/view/subjects/270705.html">270000 Biological Sciences &gt; 270700 Ecology and Evolution &gt; 270705 Palaeoecology</a><br /><a href="http://eprints.utas.edu.au/view/subjects/260112.html">260000 Earth Sciences &gt; 260100 Geology &gt; 260112 Palaeontology</a><br /><a href="http://eprints.utas.edu.au/view/subjects/260113.html">260000 Earth Sciences &gt; 260100 Geology &gt; 260113 Palynology</a></td></tr><tr><th valign="top" class="ep_row">Collections:</th><td valign="top" class="ep_row">UNSPECIFIED</td></tr><tr><th valign="top" class="ep_row">ID Code:</th><td valign="top" class="ep_row">1727</td></tr><tr><th valign="top" class="ep_row">Deposited By:</th><td valign="top" class="ep_row"><span class="ep_name_citation"><span class="person_name">dr gregory j jordan</span></span></td></tr><tr><th valign="top" class="ep_row">Deposited On:</th><td valign="top" class="ep_row">30 Aug 2007</td></tr><tr><th valign="top" class="ep_row">Last Modified:</th><td valign="top" class="ep_row">12 Feb 2008 12:29</td></tr><tr><th valign="top" class="ep_row">ePrint Statistics:</th><td valign="top" class="ep_row"><a target="ePrintStats" href="/es/index.php?action=show_detail_eprint;id=1727;">View statistics for this ePrint</a></td></tr></table><p align="right">Repository Staff Only: <a href="http://eprints.utas.edu.au/cgi/users/home?screen=EPrint::View&amp;eprintid=1727">item control page</a></p>
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